Sample Research Paper on Sulfur Deficiency in Brassica Rapa Plant



Introduction

            In plants, essential nutrients play significant roles to ensure the healthy functioning of the plants. The nutrients in plants are protein, vitamins, minerals, nucleic acids and carbohydrates. The essential nutrients are divided into macro-nutrients and micro-nutrients which have distinctive roles in plants. The macro-nutrients are calcium, sulfur, nitrogen, phosphorus, potassium and magnesium. On the other hand, micro-nutrients encompass; chlorine, manganese, iron, zinc, nickel, copper and barium (Imsande 140). When the plants are in deficient of any of the essential nutrients, their functions are affected. Some of the causes of the unavailability of the essential nutrients to the plants include; change in weather conditions, acid-like conditions and drought. The depletion of macro-nutrients in plants results in wilting decreased budding and loss of chlorophyll.  When the plant is in deficient of sulfur, growth of the plant is retarded (González-Ballester et al. 2080). Sulfur plays a vital role in the various synthesis of amino acids and in making necessary protein and hormones in plants(Rending and Oputa 433). The structure of plant cell wall is composed of several amino acids and protein, and hormones regulate plant cell wall function.  Therefore, the decreased level of sulfur in plants affects the structure of plant cell wall (Lewandowska and Sirko 465, Soetan, Olaiya and Oyewole 222). In this research, macronutrient sulfur was selected for the study since its effects and deficient in Brassica plants are easily observed and determined (Aghajanzadeh et al.  n.p).   For the aim of the experiment, the heights of separate plant groups were measured. The plant groups were arranged into three groups; one with 100% sulfur, 50% depleted of sulfur and the other 100% depleted of sulfur.  The degree of chlorosis in plants was determined using the spectrophotometer. The number of leaves was physically counted, and the root length was measured using a ruler.  Lastly, the density of the stomata was observed using a microscope.  Based on this experiment, it was hypothesized that sulfur deficiency contributed to shorter plants with fewer leaves, shorter roots, few chlorophyll pigments, and with low-density stomata. 
Methods
The nutrient solution was absorbed by fitting sixteen plant spaces from each six planting quads with a wick. Every quad was filled with soil, patted down, and refilled until each had desired quantity of the soil. In each cell, just below the surface, one seed was planted. 50 mL of the selected solution was poured per cell, and then 50 more mL of chosen solution was emptied nearby the base of the whole plant quad. Plant solutions were made distinctively for each two plant quads. In quads 1 and 2 with 32 plants in total, the prepared solution was lacking 100% sulfur. For quads 3 and 4, the solution was made to have 50% deficient in sulfur. Finally, Quads 5 and 6 were made with Hogland’s Complete Solution, which contains all the essential nutrients.
The height and root length of the plant were measured using a ruler at the end of three weeks of growing duration. The quantitative examination of chlorosis was performed using a spectrophotometer to obtain the concentration of chlorophyll in a particular leaf per plant quad. Chlorophyll in the leaf was dissolved by taking two punches of the leaf and dissolving it in 80% acetone. The obtained solution from the dissolution was grounded in the mortar using a pestle to break all the tissues into smallest particles. Due to evaporative nature of acetone, the lost amount of acetone was determined and the lost amount added to the original solution. The solution was then centrifuged at 300rpm for five minutes at 22 degree Celsius. 2ml of the resultant supernatant liquid was analyzed via spectrophotometer at four different wavelengths (470, 645, 663, 720 nm) to determine the quantities of the chlorophyll a, b and carotenoids in the designated leaf. 
The density of the stomata was obtained within areas of 1mm^2. For every treated group had one leaf to be measured. Two holes were made into the leaf, and the back of the leaf was sheltered in a thin sheet of clear nail polish. The polish dried up for 10 minutes, and the tape was put on topmost of the two holes.  The leaf impression was imprinted on the tape through clear nail polish by carefully removing the tape. The tape was then placed on a slide and observed under a microscope. Some stomata in 1x 1mm area of the leaf were counted.
Results 
Photosynthetic pigments
The average concentration of chlorophyll a in 100% sulfur, 50% sulfur treated leaf and sulfur deficient leaf were 19.936748ug/cm2, 18.356776ug/cm2, and 12.53167ug/cm2 respectively. While the chlorophyll b and carotenoid the average concentrations for the same leaves were 7.15195ug/cm2, 5.3005ug/cm2,5.2656ug/cm2 and 5.10037ug/cm2, 4.4636442ug/cm2, 3.38786812ug/cm2 (Table 1). The overall average of chlorophyll levels in three plant treatment was as shown in Fig1. 

Discussion
According to the results, only one part of the hypothesis of the study was justified. For instance, sulfur deficient in Brassica Rapa leads to plants with little chlorophyll. It was observed that the complete plant had more chlorophyll pigmentation than 50% sulfur treated plants (half) and deficient sulfur plants (Table 1, 2, 3 and 4). The concentration of chlorophyll a was high in all the Brassica Rapa plant tested, followed by chlorophyll b and then carotenoids as seen in Fig 1. The number of the leaves was more pronounced in Brassica plant in sulfur deficient, recording average mean of 6.78125(Table 6). The stomata density was high in deficient sulfur Brassica with the average mean of 65.5(Table 5). The Brassica plant which had a high concentration of sulfur was taller than its counterpart with the average mean of 22.13125. On the contrary to the hypothesis that states, “Deficient Sulfur in plants result in shorter plants," Brassica plant which was deficient in sulfur, was taller than Brassica Rapa treated with 50% sulfur solution.
In this experiment, there were negligible errors associated with the results, as can be seen in Table 3. Thus, the errors did not have a significant effect on the results.
It was identified in this study that the number of stomata density and leaves were more pronounced in sulfur deficient Brassica Rapa plant. Hence, for further study, scholars should consider carrying out research on the relationship between the number of leaves, stomata density and sulfur deficient in plants.

Works cited

Aghajanzadeh, Tahereh et al. "Atmospheric H2S And SO2 As Sulfur Source For Brassica Juncea And Brassica Rapa: Impact On The Glucosinolate Composition". Frontiers in Plant Science 6 (2015): n. pag. Web.
González-Ballester, D., Casero, D., Cokus, S., Pellegrini, M., Merchant, S. S., Grossman, A. R. 2010. RNA-Seq analysis of sulfur-deprived Chlamydomonas cells reveals aspects of acclimation critical for cell survival. American Society of Plant Biologists. 22: 2058-2084.web.
Imsande, J. 1998. Iron, sulfur, and chlorophyll deficiencies: a need for an integrative approach to plant physiology. Physiologia Plantarium. 139-144.web.
Lewandowska, Małgorzata, and Agnieszka Sirko. "Recent advances in understanding plant response to sulfur-deficiency stress." Acta Biochim Pol 55.3 (2008): 457-71.web
Rendig, V. V., Oputa, C., McComb, E. A. 1976. Effects of sulfur deficiency on non-protein nitrogen, soluble sugars, and N/S ratios in young corn plants. Department of Soils and Plant Nutrition. 423-437.web.
Soetan, K. O., C. O. Olaiya, and O. E. Oyewole. "The importance of mineral elements for humans, domestic animals and plants: A review." African Journal of Food Science 4.5 (2010): 200-222.web.

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